They appear for part of all queries, depend on language and country, and their content changes with the index.
Google says no special file, markup or technique is needed to appear in its AI features: pages are crawled and indexed by Googlebot as for normal search, and the same fundamentals apply.
How it relates to AI Mode: AI Mode is Google’s conversational search; AI Overviews are the summary inside the classic results. Search Console has announced performance reports that include both.